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Several important cellular kinases including the G-protein-coupled receptor kinases (GRKs) (Pronin et al., 2000), the casein kinase II (Sano et al., 2021), polo-like kinases (Inglis et al., 2009), and the leucine-rich repeat kinase 2 (LRRK2) (Greggio et al., 2011) mediate phosphorylation of α-synuclein